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That is the casting number which is kind of like the part number for cast items.

C5 is 1965. The rest describes more of what the part is.
 

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It is an engineering number and although it may look like a part number it is not one. If you go to your friendly Ford parts man and tell him "I need a C5OE-9425-C" he will tell you that there is no part with that number.
 
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